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Additional pathways of lysine lactoylation are β-dicarbonyl fragmentation of 1-DG isomerization product 1-deoxy-3,5-hexodiulose (Smuda et al., 2010) and the non-enzymatic acyl transfer verified from lactoyl glutathione or recently proposed from lactoyl-CoA thioester, which are controversially discussed (Varner et al., 2020)